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A theme in drupal is basically the way your site outputs information to the end user. It is made up of a few or sometimes several files that intercept the drupal output and in a sense catch that info and change it to look and act like you want it too.
This allows you to mess with the look and feel of your site without having to play with the actual hard code. Why would you want this, well it is almost impossible to destroy your website and database if your just changing the theme layer. Its like a safety and if something goes wrong picking a different them will fix most problems.
The files you will be working with will be discussed individually.